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"Only put off until tomorrow what you are willing to die having left undone"

Pablo Picasso

My latest work:

Holmes NP (unpublished) Multisensory integration: inverse effectiveness and noisy signals. Current Biology, 0:0
  [NBArticle #45652] [Cites 1]

Reader A, Tamè L, Holmes NP (unpublished) Two instances of presyncope during magnetic stimulation of the median nerve, and evaluation of resting motor threshold with transcranial magnetic stimulation (tms). , 0:0
  [NBArticle #45584]

Blanchard CCV, McGlashan HL, French B, Sperring RJ, Petrocochino B, Holmes NP (2017) Online control of prehension predicts performance on a standardised motor assessment test in 8-12 year old children. Frontiers in Psychology, 8:374
    [NBArticle #41777] [Cites 50] [CitedBy 3]

Curriculum Vitae

Positions Prizes Research Skills Symposia Teaching Supervision Courses Education Work Experience

Nicholas Paul Holmes

DOB: 24th September 1976 E-mail:

Journal Articles & Other Papers

Chapters & Books



Prizes, awards, scholarships, & fellowships:

2005-2007, Science Research Fellowship
From the Royal Commission for the Exhibition of 1851

Awarded for: The neural basis of bodily self-recognition
Held at: Espace et Action, Lyon, France, under Pr. Yves Rossetti, Head of Organisations sensori-motrice et cognitive de l'action
Year 1 Report - Year 2 Report

2005, Wellcome Trust Final Year Students Meeting, Best Poster Prize

Poster:Holmes NP, Calvert GA, Spence C (2005) Where does the body end & the world begin? Multisensory interactions in peripersonal space. Wellcome Trust Final Year Doctoral Students Meeting, London, UK, 27th January

    [NBArticle #25570]

2003, International Multisensory Research Forum, Graduate Student Symposium Award

Presentation: Holmes NP, Calvert GA, Spence C (2003) Brain, body, & space: Does tool-use *extend* visuotactile peripersonal space? International Multisensory Research Forum (IMRF), McMaster University, Canada, 16th June

    [NBArticle #9661]

2001-2005, Wellcome Trust 4-year Doctoral Prize Studentship in Neuroscience

Held at: Department of Experimental Psychology, University of Oxford, Oxford, UK, under Prof. Charles Spence, and Prof. Gemma A Calvert

Other positions:


Novartis Institute for Medical Sciences, University College, London

Running a high-throughput drug screening laboratory; Conducting a feasibility study of a model of drug absorption across the human intestine; Techniques mastered include HPLC; pharmacokinetics; cell culture; radiochemical usage; Ten thousand word report on work and findings contributed ten percent towards my degree

Scientific, research, & IT skills

Symposia organised:

Teaching experience:


Other supervision & teaching

Courses Attended:


2001-2005, 4 year PhD programme

Wellcome 4-Year Doctoral Programme in Neuroscience,The University of Oxford, Oxford, UK

2002-2005, D.Phil. in Neuroscience
Supervised by Drs Charles Spence and Gemma Calvert, respectively of the Department of Experimental Psychology and University Laboratory of Physiology, The University of Oxford

Citation: Holmes NP (2005) Of tools, mirrors, & bodies: Multisensory interactions in peripersonal space. PhD Thesis, Wolfson College, University of Oxford, 302pp.

    [NBArticle #25557] [Cites 288]
2001-2002, M.Sc. in Neuroscience
Comprising two research projects:
Holmes NP, Christensen LOD, Miall RC (2002) Reaching to a target is disrupted by repetitive transcranial magnetic stimulation (rTMS) over the cerebellum: Empirical support for state-estimation by feed-forward internal models. MSc Thesis, Wolfson College, University of Oxford, 31pp.
    [NBArticle #25559] [Cites 30]
Holmes NP, Skaliora I, Doubell TP, King AJ (2002) Sub-threshold auditory-visual interactions in the rat superior colliculus: Linearity and independence of EPSP summation. MSc Thesis, Wolfson College, University of Oxford, 35pp.
    [NBArticle #25560] [Cites 34]
See also:
Miall RC, Christensen LOD, Cain O, Stanley J (2008) Disruption of state estimation in the human lateral cerebellum. Public Library of Science Biology, 5(11):e316
      [NBArticle #25563] [CitedBy 3]
Miall RC, King D (2008) State estimation in the cerebellum. Cerebellum, 7(4):572-576
      [NBArticle #25562] [CitedBy 2]
Skaliora I, Doubell TP, Holmes NP, Nodal FR, King AJ (2004) Functional topography of converging visual and auditory inputs to neurons in the rat superior colliculus. Journal of Neurophysiology, 92(5):2933-2946
        [NBArticle #10325] [Cites 66] [CitedBy 25]

...and five lecture courses: (Methods of Analysing Neuronal Circuity; Mapping and Imaging Techniques; Computational Neuroscience; Animal Models and Clinical Aspects of Neuroscience; Molecular Neuroscience)

1996-2000, BSc in Psychology & Neuroscience

First Class B.Sc. Hons. in Psychology and Neuroscience, with a year's Industrial Experience, The University of Manchester, Manchester, UK

Third-year project:Holmes NP, Plainis S, Murray IJ (2000) Hemispheric asymmetries in perceptual-motor processing and the 'Space-Ship Plot': Simple reaction time to lateralised sinusoidal gratings. BSc Thesis, University of Manchester, 42pp.

    [NBArticle #25558] [Cites 84]

Dissertation:Holmes NP (1999) The self in its brain: Neural representations of the body. BSc Thesis, University of Manchester, 34pp.

    [NBArticle #1972] [Cites 137]

Selected course modules: Psychology: Ecological visual perception; Consciousness, philosophy, and the brain; Motor control. Neuroscience: Generation and control of movement; Neuropharmacology; Neurobiology; Neuroethology

1993-1995, A-Levels

A-Levels in Psychology, Human Biology, Maths, and AS-Statistics - all grade A; National Mathematics Contest, Silver Certificate, Exeter College, Devon

1991-1993, GCSEs

GSCEs (7A, 2B grades), including Maths and Extended Maths, Double English, Double Sciences, and two Foreign Languages, Beacon Community College, Sussex